氧化磷酸化与光合磷酸化

The electron transport system and oxidative
phosphorylation on the mitochondrial crista.
Electrons brought in from the TCA cycle by NADH are passed
along the chain of electron transport carriers.
光合磷酸化的机制 （ Photosynthesis）
绿色和紫色细菌的光合磷酸化 Properties of Microbial Photosynthetic Systems
Property Photosynthetic pigment Photosystem II Photosynthetic electron donors O2 production pattern Primary products of energy conversion Carbon source ATP Eucaryotes Chlorophyll a Present H 2O Oxygenic ATP +NADPH CO2 Cyanobacteria Green and Purple Bacteria Chlorophyll a Present H 2O Oxygenic ATP +NADPH CO2 ADP + Pi Bacteriochlorophyll Absent H2, H2S, S, Organic Matter Anoxygenic ATP Organic and/or CO2
The aerobic respiratory system of E. coli
2. 光合磷酸化 （ photosynthesis）
绿色植物与蓝细菌（Cyanobacteria）的光合色素
Chlorophylls（叶绿素） Carotenoids (类胡萝卜素) -Carotene (-胡萝卜素) Phycobiliproteins (藻胆蛋白) Phycoerythrin (藻红蛋白)
氧化磷酸化与光合磷酸化
1. 氧化磷酸化（Oxidative phosphorylation）
底物脱氢（EMP）途 径
Fructose-6-phosphate
丙酮酸
ATP
ADP
NAD+
CO2
NADH+H+
NAD+
glucose 6-磷酸葡萄糖6-磷酸葡萄糖酸
底物脱氢（HMP） 途径
乙酰磷酸
(光系统 II)
Model for the arrangement of light-harvesting chlorophylis/ bacteriochlorophylls versus reaction centers within a photosynthetic membrane. RC: reaction centers
Electron transport systems are composed of membrane associated electron carriers.
The distribution of electric potential across the
membrane drives the synthesis of ATP by ATP synthase.
NADH+H+ NAD+
5-磷酸核酮糖 NADH+H+ 5-磷酸木酮糖 3-磷酸甘油醛 Pyruvate
NADH+H+ NAD+
乙酰coA 乙醛 Ethanol
乳酸
底物脱氢（TCA） 循环
Hans Krebs Discovered the TCA cycle in 1937
He received the Nobel Prize in physiology or medicine in 1953 for his discovery of TCA cycle. He was Forced to leave Germany prior to WWII because he was Jewish.
H2S, or S or
Succinate
+ NAD+
S, or SO42-, +NADH + H&ple Nonsulfur Bacterial Photosynthesis.
Green sulfur Bacterial Photosynthesis.
Photomicrographs of phototrophic bacteria.
Photosystem I：Absorbs longer wavelength light (680nm)P700
(光系统 I)
Antennas Accessory pigments (天线色素)
Photosystem II: Traps light at shorter wavelength (680nm) P680
a: Purple bacterium; b: Green bacterium (The refractile bodies are sulfur granules deposited outside the cell)
光合磷酸化与氧化磷酸化的比较
思考题
1. 氧化磷酸化和光合磷酸化的实质是什么 2. 氧化磷酸化和光合磷酸化的重要性
A Photosynthetic Reaction Chain. The reaction center of the purple nonsulfur bacterium
Green Plant / Cyanobacteria Photosynthesis, Having two photosystems